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osed a 7 genera treatment, which was adopted by the IUCN Red List and BirdLife International, and later by the IOC in 2021, standardizing it. Occipital crest or os nuchale in Phalacrocorax carbo The cormorants and the darters have a unique bone on the back of the top of the skull known as the os nuchale or occipital style which was called a xiphoid process in early literature. This bony projection provides anchorage for the muscles that increase the force with which the lower mandible is closed. This bone and the highly developed muscles over it, the M. adductor mandibulae caput nuchale, are unique to the families Phalacrocoracidae and Anhingidae. Several evolutionary groups are still recognizable. However, combining the available evidence suggests that there has also been a great deal of convergent evolution; for example, the cliff shags are a convergent paraphyletic group. The proposed division into Phalacrocorax sensu stricto (or subfamily "Phalacrocoracinae") cormorants and Leucocarbo sensu lato (or "Leucocarboninae") shags does have some degree of merit. The resolution provided by the mtDNA 12S rRNA and ATPase subunits six and eight sequence data is not sufficient to resolve several groups to satisfaction properly; in addition, many species remain unsampled, the fossil record has not been integrated in the data, and the effects of hybridisation – known in some Pacific species especially – on the DNA sequence data are unstudied. A multigene molecular phylogenetic study published in 2014 provided a genus-level phylogeny of the family; this is now followed by most authorities, includi